9 with	a
      9 The	parameters
      9 the	model
      9 simulation	results
      9 of	sexual
      9 N	MP
      9 N	MD
      9 N	FD
      9 N	FC
      9 individual	variation
      9 from	the
      9 as	the
      9 4	0
      9 200	0
      9 0	1
      8 with	the
      8 the	social
      8 the	same
      8 the	fitness
      8 the	baseline
      8 social	cohesion
      8 public	goods
      8 period	5
      8 of	sexuality
      8 N_	mathit
      8 MP	N
      8 MP	and
      8 MD	N
      8 Journal	of
      8 fitness	of
      8 F	C
      8 and	FC
      7 to	be
      7 this	simulation
      7 the	population
      7 the	evolutionary
      7 the	altruistic
      7 simulation	are
      7 s	1
      7 population	size
      7 of	cooperators
      7 four	types
      7 can	be
      7 baseline	model
      7 are	the
      7 a	male
      7 200	200
      7 0	001
      6 ω	0
      6 μ	0
      6 κ	1
      6 Ι	1
      6 δ	0
      6 γ	0
      6 β	1
      6 α	0
      6 this	paper
      6 the	Punishment
      6 Sexuality	Game
      6 same	as
      6 Punishment	Sexuality
      6 population	of
      6 of	punishment
      6 of	I
      6 N	D
      6 N	C
      6 N	400
      6 mutation	rate
      6 is	the
      6 In	the
      6 in	an
      6 I	I
      6 d	0
      6 constant	population
      6 by	the
      6 attractiveness	The
      6 al	2011
      6 5	δ
      6 5	γ
      6 400	N
      6 1	μ
      6 1	β
      6 1	α
      6 1	s
      6 001	d
      5 Ye	et
      5 widetilde	I
      5 which	is
      5 where	the
      5 variation	in
      5 this	section
      5 the	variable
      5 the	simulation
      5 the	role
      5 The	population
      5 the	offspring
      5 the	effect
      5 society	is
      5 social	dilemma
      5 role	of
      5 robust	test
      5 represents	the
      5 probability	of
      5 parameters	in
      5 of	variable
      5 offspring	is
      5 of	each
      5 of	a
      5 MP	MD
      5 model	the
      5 mean	value
      5 mathit	MP
      5 males	and
      5 left	1
      5 it	is
      5 In	this
      5 I	into
      5 Game	with
      5 Figure	1C
      5 F	D
      5 FC	N
      5 effect	of
      5 each	individual
      5 due	to
      5 do	not
      5 and	the
      5 altruism	in
      5 6	s
     54 of	the
      5 0	0
      4 Ye	s
      4 whose	I
      4 we	have
      4 those	in
      4 the	second
      4 the	robust
      4 the	punisher
      4 the	plots
      4 theoretical	biology
      4 the	number
      4 the	mean
      4 the	exception
      4 the	behaviour
      4 that	is
      4 that	altruism
      4 surrey	ac
      4 strength	of
      4 soc	surrey
      4 Societies	and
      4 Social	Simulation
      4 s	fitness
      4 set	to
      4 results	for
      4 punishment	is
      4 proportion	of
      4 Proceedings	of
      4 population	assumption
      4 P_	mathit
      4 parental	care
      4 parameters	for
      4 of	theoretical
      4 of	sympathy
      4 of	social
      4 of	males
      4 of	cooperation
      4 of	Artificial
      4 of	an
      4 number	of
      4 MP	is
      4 MD	and
      4 mathit	FC
      4 mate	preference
      4 k	widetilde
      4 jasss	soc
      4 is	N
      4 I	right
      4 invasion	capacity
      4 in	a
      4 I	0
      4 http	jasss
      4 he	will
      4 has	been
      4 function	of
      4 for	this
      4 FD	100
      4 FC	FD
      4 FC	0
      4 existence	of
      4 exception	of
      4 e	k
      4 differences	in
      4 cooperators	and
      4 beta	left
      4 based	on
      4 attractiveness	in
      4 at	a
      4 as	those
      4 Artificial	Societies
      4 are	N
      4 and	Social
      4 and	N
      4 and	FD
      4 and	F
      4 and	D
      4 and	attractiveness
      4 an	artificial
      4 altruistic	level
      4 altruistic	behaviour
      4 altruism	is
      4 a	female
      4 ac	uk
      4 a	cost
      4 6	Ι
      4 5000	0
      4 4	ω
     42 in	the
      4 200	5000
      4 2000	200
      4 1	e
      4 1C	with
      4 0	period
      4 0	N
      3 without	sexual
      3 with	no
      3 with	Different
      3 will	suffer
      3 will	be
      3 where	N
      3 we	employ
      3 we	assume
      3 virtue	of
      3 Variable	Size
      3 types	of
      3 types	in
      3 type	of
      3 to	test
      3 to	males
      3 to	as
      3 to	a
      3 The	timing
      3 the	strength
      3 the	simulations
      3 the	results
      3 the	punishment
      3 the	proportion
      3 the	previous
      3 the	invasion
      3 the	individual
      3 the	field
      3 the	existence
      3 The	evolution
      3 the	average
      3 test	the
      3 Table	2
      3 suggest	that
      3 strategies	and
      3 society	with
      3 Size	Population
      3 serves	as
      3 sense	of
      3 selection	is
      3 second	order
      3 regardless	of
      3 referred	to
      3 rate	μ
      3 puzzle	of
      3 Punishment	and
      3 prosocial	behaviours
      3 proportional	to
      3 population	treatment
      3 population	models
      3 plots	of
      3 order	social
      3 only	males
      3 of	s
      3 of	London
      3 of	human
      3 of	fitness
      3 of	F
      3 of	both
      3 of	altruistic
      3 of	0
      3 no	sexual
      3 normal	distribution
      3 MP	The
      3 MP	s
      3 Moran	process
      3 model	The
      3 Miller	2007
      3 MD	is
      3 MD	FC
      3 mating	preference
      3 males	whose
      3 male	punishers
      3 London	B
      3 is	not
      3 is	an
      3 in	x
      3 investing	I
      3 introduction	of
      3 into	the
      3 into	altruism
      3 in	Table
      3 in	previous
      3 individuals	in
      3 individual	s
      3 increase	the
      3 in	biology
      3 I	is
      3 if	the
      3 human	society
      3 Houchmandzadeh	2015
      3 goods	game
      3 from	MP
      3 from	males
      3 form	of
      3 Finally	we
      3 Figure	7
      3 Figure	5
      3 Figure	1
      3 field	of
      3 f	I
      3 FD	is
      3 FD	200
      3 FC	and
      3 except	for
      3 evolutionary	models
      3 each	type
      3 distribution	of
      3 denotes	the
      3 D	are
      3 D	200
      3 costly	punishment
      3 concept	of
      3 computer	simulations
      3 composed	of
      3 but	it
      3 benefit	from
      3 because	the
      3 be	achieved
      3 be	a
      3 Based	on
      3 attractiveness	s
      3 at	period
      3 as	well
      3 assumption	of
      3 assume	that
      3 assumed	to
      3 are	not
      3 and	no
      3 and	females
      3 analysis	of
      3 altruistic	investment
      3 altruistic	evolution
      3 altruism	can
      3 a	costly
      3 according	to
      3 2	2
      3 200	κ
      3 19	2
      3 18564	jasss
      3 10	18564
      3 10	1016
      3 100	κ
      3 0	ω
      3 0	s
      2 μ	which
      2 γ	δ
      2 Zhejiang	University
      2 x	right
      2 with	I
      2 with	100
      2 will	punish
      2 will	favour
      2 which	will
      2 where	s
      2 where	f
      2 we	report
      2 well	as
      2 We	adopt
      2 W	D
      2 various	forms
      2 variation	and
      2 University	China
      2 Treatments	and
      2 treatment	of
      2 Traulsen	A
      2 to	MP
      2 to	model
      2 to	have
      2 to	cooperation
      2 timing	appears
      2 Thus	in
      2 the	virtue
      2 the	type
      2 the	turning
      2 the	sympathy
      2 The	strategies
      2 the	sense
      2 the	sake
      2 the	royal
      2 The	results
      2 the	puzzle
      2 the	period
      2 The	origins
      2 the	normal
      2 the	moral
      2 the	mate
      2 the	male
      2 the	majority
      2 their	genders
      2 the	investment
      2 the	introduction
      2 the	initial
      2 the	individuals
      2 the	hypothesis
      2 the	gender
      2 the	game
      2 the	four
      2 the	format
      2 the	extended
      2 the	emergence
      2 the	egoists
      2 the	driving
      2 the	distribution
      2 the	differences
      2 the	definition
      2 the	contribution
      2 the	benefit
      2 the	Baseline
      2 the	assumption
      2 the	analysis
      2 that	MP
      2 that	altruistic
      2 than	the
      2 test	on
      2 Table	1
      2 sympathy	in
      2 sum	f
      2 suffer	from
      2 such	as
      2 solve	the
      2 society	will
      2 society	The
      2 society	of
      2 society	as
      2 social	norms
      2 social	equilibrium
      2 s	model
      2 Smith	E
      2 simulations	are
      2 Simulation	Results
      2 simulation	in
      2 simplicity	we
      2 shows	that
      2 should	be
      2 shapes	the
      2 Sexual	selection
      2 sexual	reproduction
      2 sexually	attractive
      2 sexuality	in
      2 sexuality	as
      2 sexuality	and
      2 Sexual	Attractiveness
      2 settings	in
      2 selection	in
      2 selection	can
      2 selection	and
      2 Science	316
      2 sake	of
      2 royal	society
      2 right	mathit
      2 right	delta
      2 review	of
      2 results	in
      2 resource	limitations
      2 report	the
      2 repeated	computer
      2 reciprocal	altruism
      2 reasonable	to
      2 quality	and
      2 Punishment	without
      2 punishment	The
      2 punishment	on
      2 punishment	in
      2 punishment	from
      2 punish	individuals
      2 punishers	and
      2 provide	a
      2 process	of
      2 process	is
      2 probability	μ
      2 previous	sections
      2 previous	literatures
      2 Preston	2013
      2 present	in
      2 preference	to
      2 population	payoff
      2 population	model
      2 population	has
      2 population	dynamics
      2 population	distribution
      2 population	and
      2 plays	the
      2 periods	The
      2 penduline	tits
      2 paper	we
      2 paper	is
      2 own	fitness
      2 over	parental
      2 overlap	with
      2 overcome	the
      2 our	model
      2 Other	articles
      2 origins	of
      2 on	this
      2 one	of
      2 on	each
      2 on	altruism
      2 of	γ
      2 of	virtue
      2 of	their
      2 of	punishers
      2 of	prosocial
      2 of	N
      2 of	MP
      2 of	morality
      2 of	mating
      2 of	male
      2 of	justice
      2 of	individuals
      2 of	individual
      2 of	high
      2 of	females
      2 of	FC
      2 of	economics
      2 of	death
      2 of	costly
      2 of	cooperative
      2 of	birth
      2 of	biology
      2 of	being
      2 numerical	simulation
      2 Nowak	M
      2 not	significant
      2 Note	that
      2 note	that
      2 noted	that
      2 No	punishment
      2 no	punishment
      2 N	is
      2 nearly	no
      2 nearly	every
      2 MP	N_
      2 more	than
      2 more	cooperative
      2 moral	virtues
      2 morality	in
      2 model	where
      2 models	with
      2 models	in
      2 models	based
      2 model	of
      2 model	can
      2 model	and
      2 might	be
      2 mechanism	of
      2 mean	altruistic
      2 McAndrew	2002
      2 Mating	Preference
      2 mathit	MD
      2 mathit	FD
      2 mathit	alpha
      2 male	with
      2 male	sum
      2 males	females
      2 M	A
      2 limited	resources
      2 left	N_
      2 lead	to
      2 leads	to
      2 Koella	2000
      2 journal	of
      2 J	C
      2 J	2011
      2 It	should
      2 is	that
      2 is	set
      2 is	referred
      2 is	reasonable
      2 is	proportional
      2 is	identical
      2 is	designed
      2 is	composed
      2 is	assumed
      2 is	altruistic
      2 is	advantageous
      2 In	Ye
      2 invest	1
      2 in	various
      2 in	turn
      2 in	simulation
      2 In	other
      2 in	MP
      2 in	models
      2 initial	population
      2 Individual	variation
      2 individual	in
      2 increase	in
      2 in	altruistic
      2 in	altruism
      2 imply	that
      2 I	male
      2 I	in
      2 identical	to
      2 I	ast
      2 hypothesis	is
      2 human	altruism
      2 html	doi
      2 However	the
      2 Houchmandzadeh	s
      2 his	probability
      2 high	altruistic
      2 have	been
      2 Hauert	et
      2 has	a
      2 Hang	Ye
      2 Hamilton	1981
      2 Hamilton	1964
      2 GINTIS	H
      2 gender	based
      2 game	of
      2 from	FC
      2 For	the
      2 form	for
      2 format	of
      2 For	each
      2 For	authors
      2 F	J
      2 Figure	9
      2 Figure	6
      2 Figure	2
      2 Figure	1A
      2 female	will
      2 females	The
      2 females	investing
      2 females	invest
      2 females	in
      2 female	cooperators
      2 f	e
      2 FD	are
      2 FC	is
      2 FC	in
      2 FC	are
      2 favour	males
      2 e	ωP
      2 Extension	to
      2 extended	models
      2 explain	the
      2 Evolution	of
      2 evolutionary	model
      2 evolutionary	equilibrium
      2 e	the
      2 equilibrium	where
      2 employ	the
      2 e	MP
      2 emergence	of
      2 e	g
      2 effects	of
      2 each	period
      2 E	A
      2 dynamic	evolutionary
      2 driving	force
      2 dominance	over
      2 does	not
      2 distribution	in
      2 different	treatments
      2 difference	between
      2 designed	to
      2 delta	N_
      2 definition	of
      2 defined	in
      2 c	which
      2 C	Traulsen
      2 C	Punishment
      2 cooperators	who
      2 cooperative	society
      2 cooperative	punishment
      2 cooperation	Science
      2 contribution	of
      2 conflict	over
      2 conducted	by
      2 computer	simulation
      2 cohesion	can
      2 care	when
      2 CAPTION	Table
      2 capacity	of
      2 can	increase
      2 can	explain
      2 by	Hauert
      2 B	Punishment
      2 BOYD	R
      2 both	punishment
      2 biology	and
      2 biological	sciences
      2 be	noted
      2 benefit	the
      2 benefits	from
      2 being	an
      2 being	altruistic
      2 behaviours	of
      2 behaviour	based
      2 B	biological
      2 Baseline	Model
      2 based	sexual
      2 B2	and
      2 AXELROD	R
      2 Axelrod	Hamilton
      2 average	population
      2 attractiveness	via
      2 attractiveness	variable
      2 attractiveness	N
      2 attractiveness	from
      2 attractiveness	and
      2 a	superlative
      2 a	strong
      2 ast	male
      2 as	punishers
      2 as	in
      2 a	sexually
      2 artificial	society
      2 artificial	life
      2 a	particular
      2 A	No
      2 an	MP
      2 an	isolated
      2 an	individual
      2 an	increasing
      2 an	FC
      2 a	net
      2 and	thus
      2 and	social
      2 and	population
      2 and	only
      2 and	numerical
      2 and	individual
      2 and	human
      2 and	genetics
      2 and	defectors
      2 and	costly
      2 and	cooperation
      2 and	behaviours
      2 and	B3
      2 and	admiration
      2 a	mean
      2 altruistic	is
      2 altruistic	cost
      2 altruism	the
      2 altruism	as
      2 altruism	and
      2 alpha	I
      2 a	higher
      2 a	generalised
      2 a	function
      2 a	dynamic
      2 advantage	of
      2 adopt	the
      2 admiration	from
      2 a	constant
      2 achieved	in
      2 achieved	by
      2 A3	0
      2 8	D
      2 8	B
     27 sexual	attractiveness
      2 7	Figure
      2 50	periods
      2 4	and
      2 4	20
     23 evolution	of
      2 3	4
      2 3	2
      2 316	5833
     22 to	the
     22 of	altruism
      2 2	http
      2 2	html
      2 2012	In
      2 2011	The
      2 1	with
      2 1	The
      2 1	period
      2 1	and
      2 18	3
      2 14	4
      2 13	3
      2 1	2000
      2 1016	j
      2 10	1098
      2 100	Defectors
      2 1000	200
      2 1	000
      2 0	Ι
     20 for	the
      2 0	8
      2 0	7
      2 0	2000
      2 0	200
      2 0	1000
      2 000	C
      2 000	B
      2 000	and
      1 ωP	where
      1 ωP	Removed
      1 ω	denotes
      1 ω	1
      1 μ	The
      1 μ	nvest
      1 μ	Initially
      1 μ	has
      1 κ	are
      1 δ	to
      1 δ	For
      1 δ	c
      1 δ	Attractivenesss
      1 γ	and
      1 β	and
      1 αI	However
      1 Zhun	Deng
      1 Zeng	and
      1 Zaballa	L
      1 Zaballa	2010
      1 Z	2016
      1 your	browser
      1 Yize	Chen
      1 Yet	we
      1 yet	in
      1 YE	H
      1 Ye	b
      1 Y	Chen
      1 Y	2011
      1 xN_C	6
      1 x	is
      1 x	increases
      1 x	in
      1 x	Houchmandzadeh
      1 x	from
      1 x	frac
      1 x	following
      1 x	at
      1 x	are
      1 W	Weissing
      1 W	Smith
      1 would	be
      1 world	if
      1 work	was
      1 works	have
      1 words	F
      1 women	s
      1 with	viscous
      1 with	three
      1 with	this
      1 with	these
      1 With	the
      1 withstands	the
      1 with	probability
      1 Without	the
      1 without	the
      1 without	exploring
      1 without	attractiveness
      1 with	others
      1 with	more
      1 with	Mating
      1 with	Javascript
      1 within	the
      1 with	individual
      1 with	Houchmandzadeh
      1 withhold	long
      1 with	hard
      1 with	four
      1 with	females
      1 with	F
      1 with	cellular
      1 with	altruistic
      1 Wilson	D
      1 Wilson	1999
      1 will	persistently
      1 will	obtain
      1 will	not
      1 will	never
      1 will	modify
      1 will	look
      1 will	lead
      1 will	invest
      1 will	incur
      1 will	increase
      1 will	express
      1 wild	type
      1 widen	the
      1 whom	the
      1 whole	society
      1 whole	a
      1 who	do
      1 who	admire
      1 while	the
      1 while	sexuality
      1 while	only
      1 while	mate
      1 while	FC
      1 while	decreasing
      1 which	means
      1 which	may
      1 which	greatly
      1 which	fits
      1 whether	the
      1 where	prosocial
      1 where	population
      1 where	players
      1 where	P
      1 where	mating
      1 where	I
      1 where	everyone
      1 whereas	MD
      1 where	altruists
      1 When	the
      1 when	the
      1 when	punishment
      1 when	N
      1 when	individuals
      1 when	females
      1 when	composed
      1 when	care
      1 when	both
      1 we	will
      1 we	utilize
      1 We	then
      1 we	speculate
      1 We	select
      1 we	provide
      1 we	propose
      1 we	plot
      1 We	perform
      1 We	now
      1 We	model
      1 well	Now
      1 well	fitted
      1 Weissing	F
      1 we	illustrate
      1 We	found
      1 we	do
      1 we	discuss
      1 we	compare
      1 we	classify
      1 We	believe
      1 We	are
      1 we	apply
      1 we	adopt
      1 we	acknowledge
      1 way	to
      1 ways	of
      1 ways	1
      1 way	for
      1 way	Barta
      1 was	supported
      1 was	set
      1 was	partially
      1 was	contingent
      1 viscous	populations
      1 virtue	To
      1 virtues	The
      1 virtues	can
      1 virtues	biological
      1 virtue	Penguin
      1 virtue	mutualism
      1 violation	Potgieter
      1 violation	of
      1 viewed	Most
      1 viewed	as
      1 via	various
      1 via	the
      1 via	sexual
      1 via	methods
      1 via	inhibition
      1 Via	freedom
      1 via	a
      1 versus	the
      1 Varying	population
      1 varying	population
      1 various	timings
      1 various	selections
      1 various	parameters
      1 various	disciplines
      1 various	adjusted
      1 Variation	Variations
      1 variation	to
      1 Variations	in
      1 variation	behind
      1 variation	assumption
      1 variation	appears
      1 variables	such
      1 VAN	DIJK
      1 Van	Dijk
      1 Value	of
      1 value	larger
      1 value	from
      1 value	altruism
      1 utilize	their
      1 usually	associated
      1 useful	suggestions
      1 up	the
      1 upon	the
      1 Unto	others
      1 unselfish	behavior
      1 University	Press
      1 unit	of
      1 unique	perspective
      1 uninvolved	bystanders
      1 under	variable
      1 under	the
      1 understood	differently
      1 understanding	of
      1 unchanged	the
      1 UK	SHUTTERS
      1 uk	19
      1 uk	18
      1 uk	14
      1 uk	13
      1 typical	public
      1 typically	ignore
      1 type	within
      1 type	that
      1 types	To
      1 types	i
      1 types	cooperator
      1 type	i
      1 type	Calculate
      1 type	are
      1 two	ways
      1 two	steps
      1 T	W
      1 Tu	Yize
      1 turn	provide
      1 turning	points
      1 turning	point
      1 turn	he
      1 turn	enhance
      1 trying	to
      1 TRIVERS	R
      1 Trivers	1971
      1 tribute	to
      1 tribes	Keywords
      1 treatments	Figure
      1 treatments	as
      1 treatment	offers
      1 treatment	is
      1 treatment	in
      1 treated	as
      1 Traulsen	et
      1 TRAULSEN	A
      1 Traulsen	2005
      1 Trans	R
      1 Transaction	Publishers
      1 traits	may
      1 trait	but
      1 tragedy	and
      1 tradition	from
      1 traditional	culture
      1 traditional	altruistic
      1 to	Zhun
      1 to	whom
      1 to	Variable
      1 to	value
      1 to	type
      1 to	themselves
      1 total	population
      1 to	suppose
      1 To	summary
      1 to	submit
      1 to	study
      1 To	solve
      1 to	solve
      1 To	simplify
      1 to	sexual
      1 to	sex
      1 to	see
      1 to	satisfy
      1 to	resolve
      1 to	report
      1 to	replace
      1 to	rename
      1 to	redistribute
      1 to	question
      1 to	punish
      1 to	public
      1 to	provide
      1 to	produce
      1 to	pay
      1 to	overcome
      1 to	offer
      1 Tooby	J
      1 to	note
      1 To	keep
      1 to	invest
      1 To	introduce
      1 to	Individual
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      1 to	Hamilton
      1 to	generate
      1 to	fertilization
      1 To	extract
      1 to	explain
      1 To	expand
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      1 to	each
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      1 to	coordinate
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      1 To	clarify
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      1 to	benefit
      1 to	B3
      1 to	B2
      1 to	another
      1 to	altruism
      1 to	advertise
      1 to	abundant
      1 to	1
      1 to	0
      1 T	Komdeur
      1 tits	Proceedings
      1 tits	has
      1 timings	of
      1 timing	of
      1 timing	does
      1 thus	there
      1 thus	increase
      1 through	two
      1 through	the
      1 three	types
      1 three	simulations
      1 three	repeated
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      1 This	work
      1 This	violation
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      1 This	page
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      1 this	assumption
      1 this	approach
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      1 They	share
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      1 they	do
      1 the	wild
      1 the	whole
      1 the	way
      1 the	various
      1 The	variable
      1 The	value
      1 The	treatment
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      1 the	traditional
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      1 The	study
      1 The	strength
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      1 The	spatial
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      1 The	simulation
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      1 the	Shiwiar
      1 the	sexual
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      1 these	simulations
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      1 these	authors
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      1 the	s
      1 the	Royal
      1 The	result
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      1 the	resolution
      1 Therefore	an
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      1 the	rational
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      1 the	public
      1 the	proposed
      1 The	process
      1 the	process
      1 The	probability
      1 the	preference
      1 the	predecessors
      1 the	preceding
      1 the	power
      1 The	possible
      1 The	possibility
      1 The	plots
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      1 the	player
      1 the	perspective
      1 the	persistent
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      1 the	payoff
      1 the	particular
      1 The	outcome
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      1 theory	social
      1 theory	Gintis
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      1 theoretical	biologists
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      1 the	nonparticipant
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      1 then	her
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      1 the	next
      1 the	nemesis
      1 the	National
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      1 the	names
      1 The	mutation
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      1 themselves	and
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      1 the	models
      1 the	midpoint
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      1 The	mechanism
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      1 The	mate
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      1 the	linear
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      1 the	limit
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      1 the	later
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      1 their	sexual
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      1 their	own
      1 their	generation
      1 The	investment
      1 The	introduction
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      1 the	interaction
      1 The	initial
      1 The	increase
      1 the	incentives
      1 the	implicit
      1 The	ignorance
      1 the	I
      1 The	hypothesis
      1 the	human
      1 the	historical
      1 the	highest
      1 The	hierarchy
      1 the	group
      1 the	given
      1 The	genetical
      1 the	gap
      1 The	fundamental
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      1 the	formula
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      1 the	force
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      1 the	fact
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      1 the	exponential
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      1 the	exact
      1 The	Evolution
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      1 the	effects
      1 The	dynamics
      1 The	dynamic
      1 the	dynamic
      1 the	dominant
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      1 The	difference
      1 The	dependency
      1 the	defectors
      1 the	dead
      1 the	Darwinian
      1 the	cost
      1 the	corresponding
      1 the	cooperator
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      1 the	concurrent
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      1 The	computer
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      1 the	co
      1 the	change
      1 The	case
      1 the	case
      1 the	carrying
      1 the	Cambridge
      1 the	calculations
      1 the	breadth
      1 The	Blessing
      1 The	biology
      1 the	benefits
      1 The	benefit
      1 The	basic
      1 the	balance
      1 the	assumptions
      1 The	apparent
      1 the	altruists
      1 the	altruist
      1 The	altruism
      1 the	agent
      1 The	advantage
      1 the	advantage
      1 the	admiration
      1 the	adjusted
      1 the	actor
      1 the	accuracy
      1 The	above
      1 that	will
      1 that	when
      1 that	was
      1 that	varying
      1 that	social
      1 that	report
      1 that	our
      1 that	only
      1 that	mutation
      1 that	moral
      1 that	models
      1 that	it
      1 that	inter
      1 that	increases
      1 that	in
      1 that	I
      1 that	FD
      1 that	cooperation
      1 that	can
      1 that	benefits
      1 that	any
      1 that	an
      1 that	a
      1 than	in
      1 than	him
      1 than	her
      1 than	four
      1 than	200
      1 than	10
      1 than	0
      1 T	Hales
      1 test	with
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      1 test	for
      1 term	love
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      1 T	E
      1 Tanzhe	Tang
      1 Tang	a
      1 Tan	F
      1 talked	about
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      1 take	power
      1 take	part
      1 taken	into
      1 taken	alone
      1 take	dominance
      1 T	2002
      1 Székely	T
      1 systems	Transaction
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      1 sympathy	The
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      1 sympathy	proposed
      1 sympathy	individuals
      1 sympathy	can
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      1 switch	to
      1 survive	under
      1 Surprisingly	the
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      1 superlative	example
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      1 super	heroines
      1 superheroes	and
      1 summary	a
      1 sum	I_i
      1 SUGIYAMA	L
      1 Sugiyama	et
      1 suggestions	we
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      1 suffers	nearly
      1 suffer	punishment
      1 success	Alonzo
      1 substructure	of
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      1 Studying	artificial
      1 studies	In
      1 studied	the
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      1 structure	may
      1 structure	i
      1 Structured	Societies
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      1 strong	stabilizer
      1 strong	sense
      1 Strong	reciprocity
      1 strict	stereotypes
      1 strict	constraint
      1 Strengths	of
      1 strength	0
      1 strategy	Ye
      1 strategy	that
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      1 strategy	in
      1 strategy	i
      1 strategies	Journal
      1 S	Tooby
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      1 steps	of
      1 steps	1
      1 step	of
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      1 Statistics	Statistics
      1 Statistics	Most
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      1 stabilizer	for
      1 stability	of
      1 S	T
      1 s	sympathy
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      1 s	simulation
      1 s	sexual
      1 Springer	Science
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      1 spatial	spread
      1 spatial	distribution
      1 s	own
      1 South	African
      1 sources	to
      1 some	initial
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      1 solved	by
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      1 solace	or
      1 society	whereas
      1 society	where
      1 society	when
      1 society	structure
      1 society	Overall
      1 society	Otherwise
      1 Society	of
      1 society	N
      1 society	may
      1 society	into
      1 society	i
      1 society	he
      1 society	given
      1 society	from
      1 society	Finally
      1 society	Figure
      1 society	compete
      1 society	Besides
      1 society	are
      1 society	and
      1 society	although
      1 society	54
      1 societies	with
      1 societies	typically
      1 Societies	Journal
      1 societies	In
      1 societies	in
      1 social	structures
      1 social	selection
      1 social	scrutiny
      1 Social	Science
      1 social	justice
      1 sociality	Journal
      1 social	exchange
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      1 social	cooperation
      1 social	behavior
      1 social	altruistic
      1 Soc	B
      1 so	called
      1 Sober	Wilson
      1 SOBER	E
      1 s	Note
      1 sN_	mathit
      1 Smith	2001
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      1 size	on
      1 size	N
      1 size	is
      1 size	in
      1 size	due
      1 size	benefits
      1 size	and
      1 s	It
      1 s	is
      1 s	investments
      1 single	period
      1 since	a
      1 s	In
      1 simulation	where
      1 simulation	We
      1 simulations	where
      1 simulations	we
      1 simulations	that
      1 simulations	of
      1 simulations	in
      1 simulations	However
      1 Simulations	have
      1 simulations	have
      1 simulations	Hang
      1 simulations	Figure
      1 simulations	due
      1 simulation	requires
      1 simulation	practices
      1 simulation	models
      1 simulation	model
      1 simulation	is
      1 simulation	indicates
      1 Simulation	19
      1 Simulation	18
      1 Simulation	14
      1 Simulation	13
      1 simulated	model
      1 simplify	the
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      1 simple	assumptions
      1 similar	to
      1 similar	Our
      1 significant	as
      1 significant	and
      1 significant	amplifier
      1 Signature	in
      1 signal	to
      1 signalling	theory
      1 signaling	theories
      1 signaling	in
      1 Signaling	and
      1 Sigmund	K
      1 s	i
      1 SHUTTERS	S
      1 Shutters	Hales
      1 show	the
      1 show	that
      1 shows	the
      1 shown	in
      1 show	favour
      1 showed	that
      1 Shoresh	N
      1 Shiwiar	of
      1 She	will
      1 she	will
      1 She	also
      1 share	the
      1 shares	the
      1 share	nearly
      1 shape	the
      1 S	H
      1 s	genes
      1 s	Figure
      1 s	favour
      1 sexual	partners
      1 sexuality	to
      1 sexuality	shapes
      1 sexuality	possibly
      1 sexuality	is
      1 Sexuality	has
      1 Sexuality	Evolution
      1 sexuality	Baseline
      1 sexuality	__________________________________________________________________
      1 sexual	capital
      1 Sexual	attractiveness
      1 sexual	approach
      1 sex	provides
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      1 setup	We
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      1 setting	The
      1 settings	to
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      1 Set	the
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      1 separate	and
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      1 self	reinforcing
      1 selfless	help
      1 self	consciousness
      1 select	several
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      1 Selection	the
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      1 selections	at
      1 selection	of
      1 selection	models
      1 selection	may
      1 Selection	in
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      1 selection	Figure
      1 selection	favours
      1 selection	as
      1 selection	an
      1 selection	among
      1 selecting	them
      1 see	the
      1 see	Figure
      1 section	we
      1 section	the
      1 sections	The
      1 sections	is
      1 sections	In
      1 sections	except
      1 section	if
      1 section	FC
      1 section	except
      1 section	Discussion
      1 section	at
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      1 second	step
      1 S	D
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      1 s	cooperative
      1 s	contribution
      1 sciences	rspb20112297
      1 sciences	99
      1 Sciences	279
      1 sciences	267
      1 Science	Fund
      1 Science	Business
      1 Science	327
      1 Science	211
      1 science	1183068
      1 science	11
      1 Scelza	B
      1 scale	of
      1 scale	N
      1 scaled	the
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      1 Save	citation
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      1 s	asexual
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      1 salient	However
      1 saints	of
      1 sacrifice	their
      1 s	2015
      1 S	2007
      1 S	2002
      1 S	2001
      1 s	2000
      1 S	1999
      1 S0305004100033193	NEFF
      1 S0022	5193
      1 runs	synchronously
      1 rstb	2015
      1 rss	JASSS
      1 rspb20112297	YE
      1 rspb	2000
      1 R	Soc
      1 R	Richerson
      1 Royal	Society
      1 routines	Figure
      1 robust	tests
      1 robustness	checks
      1 Robustness	Check
      1 road	to
      1 R	Mathew
      1 R	M
      1 R	L
      1 right	sN_
      1 right	N_D
      1 right	left
      1 right	I
      1 RIDLEY	M
      1 Ridley	1997
      1 Richerson	P
      1 Richerson	1990
      1 R	Hamilton
      1 rewards	A
      1 revolution	trying
      1 review	E
      1 results	withhold
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      1 Results	The
      1 results	The
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      1 results	No
      1 results	In
      1 results	imply
      1 Results	for
      1 results	describing
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      1 result	coincides
      1 response	of
      1 respectively	x
      1 respectively	The
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      1 respectively	Figure
      1 respectively	2
      1 resources	Extension
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      1 Researches	in
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      1 religions	social
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      1 related	to
      1 reinforcing	process
      1 reinforce	the
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      1 References	ALEXANDER
      1 referees	We
      1 redistribute	the
      1 reciprocity	and
      1 recent	researches
      1 receives	the
      1 receives	admiration
      1 receive	nearly
      1 receive	female
      1 Received	17
      1 real	world
      1 reality	better
      1 realise	that
      1 Read	in
      1 reach	the
      1 reaches	its
      1 R	E
      1 R	D
      1 R	B
      1 rational	consumption
      1 rate	The
      1 rate	Simulation
      1 rate	can
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      1 Random	processes
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      1 raises	the
      1 RAIHANI	N
      1 Raihani	et
      1 R	2010
      1 R	1984
      1 quite	similar
      1 quite	optimistic
      1 question	the
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      1 quarterly	review
      1 qualified	as
      1 pure	altruism
      1 punish	the
      1 punish	or
      1 punish	only
      1 Punishment	γ
      1 punishment	without
      1 punishment	that
      1 punishment	strategy
      1 punishment	social
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      1 punishment	Science
      1 punishment	model
      1 punishment	mechanism
      1 Punishment	Mating
      1 punishment	game
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      1 punishment	favours
      1 punishment	evolution
      1 punishment	cements
      1 punishment	can
      1 punishment	called
      1 punishment	but
      1 punishment	Boyd
      1 Punishment	Attractiveness
      1 punishment	are
      1 punish	MD
      1 punishes	defectors
      1 punisher	to
      1 punisher	They
      1 punishers	Ye
      1 punishers	will
      1 punishers	Sexual
      1 punishers	In
      1 punishers	in
      1 Punishers	impose
      1 punishers	from
      1 punishers	cooperators
      1 Punishers	benefit
      1 punishers	Actually
      1 punisher	s
      1 Punisher	MD
      1 punisher	In
      1 punisher	directly
      1 punisher	and
      1 punish	defectors
      1 Publishers	ALONZO
      1 Published	31
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      1 psychology	of
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      1 Psychological	bulletin
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      1 process	The
      1 process	Physical
      1 process	permits
      1 process	Moran
      1 process	follows
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      1 problem	in
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      1 probability	d
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      1 Press	SUGIYAMA
      1 press	Current
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      1 Preference	Sexual
      1 preference	Religions
      1 preference	or
      1 preference	on
      1 preference	is
      1 Preference	Download
      1 preference	altruistic
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      1 predecessors	of
      1 preceding	sections
      1 praise	solace
      1 practices	in
      1 pp	60
      1 power	to
      1 power	of
      1 power	in
      1 POTGIETER	F
      1 Potgieter	2011
      1 possibly	plays
      1 possibly	invade
      1 possible	growth
      1 possibility	of
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      1 population	unchanged
      1 Population	The
      1 population	the
      1 population	structure
      1 populations	Koella
      1 Population	size
      1 populations	Exact
      1 population	reaches
      1 Population	N
      1 population	is
      1 population	induced
      1 population	in
      1 population	frequency
      1 population	denoted
      1 population	could
      1 population	constraint
      1 population	condition
      1 Population	Assumptions
      1 Population	Assumption
      1 population	as
      1 population	All
      1 population	A1
      1 Population	A
      1 population	A
      1 Pop	is
      1 Pop	Dynamics
      1 politics	also
      1 point	where
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      1 P	MP
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      1 plot	of
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      1 players	With
      1 players	could
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      1 Pitcher	T
      1 Pitcher	2005
      1 pioneers	we
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      1 Physical	review
      1 Physica	D
      1 Phil	Trans
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      1 Philosophical	Analysis
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      1 perspective	to
      1 perspectives	on
      1 perspectives	Finally
      1 perspective	serves
      1 perspective	of
      1 perspective	altruistic
      1 persistent	puzzle
      1 persistently	dominate
      1 permit	the
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      1 period	To
      1 period	The
      1 periods	We
      1 Periods	The
      1 periods	However
      1 periods	Figure
      1 periods	CAPTION
      1 periods	C
      1 periods	B
      1 period	say
      1 Period	Punishment
      1 period	less
      1 period	i
      1 period	For
      1 period	as
      1 period	are
      1 period	Additionally
      1 Period	5
      1 period	100
      1 period	0
      1 perform	robustness
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      1 Penguin	UK
      1 PDF	Tanzhe
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      1 P	Calculate
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      1 partial	substitute
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      1 parameters	are
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      1 P	1958
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      1 Our	simple
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      1 others	Gintis
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      1 other	moral
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      1 other	cooperators
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      1 or	more
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      1 or	equivalently
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      1 or	defectors
      1 or	cooperator
      1 or	artificial
      1 or	admire
      1 opt	out
      1 optimistic	in
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      1 opens	up
      1 on	various
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      1 on	other
      1 on	MD
      1 only	the
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      1 only	in
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      1 only	care
      1 on	high
      1 on	defectors
      1 on	cooperators
      1 once	again
      1 on	altruistic
      1 of	μ
      1 of	δ
      1 of	x
      1 of	virtues
      1 of	unselfish
      1 of	society
      1 of	simulation
      1 of	simplicity
      1 of	Sexuality
      1 of	Sexual
      1 of	sciences
      1 of	revolution
      1 of	reciprocal
      1 of	pure
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      1 of	public
      1 of	psychology
      1 of	Population
      1 of	population
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      1 of	periods
      1 of	people
      1 of	penduline
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      1 of	other
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      1 of	newborn
      1 of	near
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      1 of	mutation
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      1 of	mathematical
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      1 of	limited
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      1 of	having
      1 of	genetic
      1 of	generations
      1 of	genders
      1 of	game
      1 offsprings	is
      1 offspring	care
      1 of	four
      1 of	fixed
      1 of	Figure
      1 offers	a
      1 offer	public
      1 of	female
      1 of	FD
      1 of	evolutionary
      1 of	evolution
      1 of	egoists
      1 of	education
      1 of	edition
      1 of	Ecuadorian
      1 of	Economics
      1 of	ecological
      1 of	determining
      1 of	defector
      1 of	Cooperation
      1 of	constant
      1 of	conflict
      1 of	computer
      1 of	cognitive
      1 of	causality
      1 of	c
      1 of	attractiveness
      1 of	asexual
      1 of	any
      1 of	another
      1 of	Altruism
      1 of	agents
      1 of	A1
      1 of	50
      1 Oct	2015
      1 obtain	a
      1 observed	in
      1 observations	and
      1 N	β
      1 N	x
      1 N	we
      1 nvest	punish
      1 numbers	of
      1 number	13AZD061
      1 Now	we
      1 now	The
      1 now	conduct
      1 now	an
      1 novel	sources
      1 not	the
      1 not	talked
      1 not	suggest
      1 not	selecting
      1 not	punish
      1 not	necessarily
      1 not	model
      1 nothing	to
      1 not	elucidate
      1 noted	by
      1 not	contribute
      1 not	complete
      1 not	clear
      1 not	appear
      1 not	an
      1 not	admire
      1 notable	Figure
      1 no	strict
      1 no	self
      1 norm	violation
      1 norms	and
      1 norms	also
      1 No	remarkable
      1 non	punishment
      1 nonparticipation	Jaffe
      1 nonparticipant	strategy
      1 nonlinear	phenomena
      1 Non	Gaussian
      1 no	MP
      1 no	more
      1 no	genetic
      1 no	fluctuations
      1 no	attractiveness
      1 no	altruistic
      1 no	admiration
      1 N	Nowak
      1 N	male
      1 N	left
      1 N	J
      1 N	frac
      1 N	female
      1 next	sections
      1 next	period
      1 new	form
      1 New	evolutionary
      1 newborns	equals
      1 newborn	offsprings
      1 newborn	according
      1 never	realise
      1 net	loss
      1 net	increase
      1 nemesis	of
      1 neglected	and
      1 Neff	Pitcher
      1 NEFF	B
      1 necessary	to
      1 necessarily	imply
      1 nearly	the
      1 Nearly	no
      1 near	extinction
      1 N_D	xN_C
      1 nature	randomly
      1 Nature	415
      1 National	Social
      1 national	academy
      1 narrow	way
      1 narrow	road
      1 names	of
      1 names	in
      1 N	7
      1 N	5
      1 mutualism	altruism
      1 mutation	plays
      1 mutation	changing
      1 Mutant	with
      1 mutant	i
      1 Multilevel	selection
      1 MP	with
      1 MP	will
      1 MP	while
      1 MP	receives
      1 MP	P
      1 MP	Male
      1 MP	beta
      1 MP	also
      1 MP	4
      1 MP	2
      1 motivation	to
      1 motivation	for
      1 Most	viewed
      1 most	males
      1 most	grateful
      1 most	females
      1 Most	cited
      1 Moreover	a
      1 more	individuals
      1 more	he
      1 more	appropriate
      1 more	altruistic
      1 Moran	Process
      1 MORAN	P
      1 Moran	1958
      1 moral	traits
      1 moral	systems
      1 moral	sense
      1 moral	psychologists
      1 moral	philosophers
      1 Morality	as
      1 moral	concept
      1 molecular	Langton
      1 Molecular	Ecology
      1 modify	this
      1 modern	civilisations
      1 model	Ye
      1 model	with
      1 model	We
      1 model	we
      1 model	Varying
      1 models	where
      1 models	when
      1 models	we
      1 models	the
      1 models	showed
      1 models	settings
      1 models	involving
      1 model	since
      1 model	setups
      1 Model	setup
      1 Model	settings
      1 Model	section
      1 models	are
      1 models	and
      1 models	A2
      1 model	operates
      1 model	opens
      1 model	MP
      1 model	Model
      1 model	loses
      1 modelling	and
      1 model	is
      1 model	Initialize
      1 Model	In
      1 model	in
      1 model	human
      1 model	has
      1 model	Figure
      1 model	dynamics
      1 model	Additionally
      1 model	2015
      1 M	Messick
      1 M	Jia
      1 MILLER	G
      1 midpoint	I
      1 methods	of
      1 method	of
      1 Messick	D
      1 Messick	2000
      1 mentioned	before
      1 member	of
      1 Medicine	Zhejiang
      1 Media	BIRD
      1 mechanism	runs
      1 mechanism	for
      1 measure	of
      1 Meanwhile	the
      1 Mean	Value
      1 means	that
      1 mean	of
      1 MD	Thus
      1 MD	takes
      1 MD	suffers
      1 MD	N_
      1 MD	Male
      1 MD	except
      1 MD	by
      1 MD	beta
      1 MCANDREW	F
      1 may	serve
      1 may	possibly
      1 may	lead
      1 may	in
      1 may	disfavour
      1 may	be
      1 may	argue
      1 MATLAB	codes
      1 Mathew	S
      1 Mathew	2007
      1 Mathematical	proceedings
      1 mathematical	biology
      1 Mate	preference
      1 mate	choice
      1 Martu	women
      1 Mar	2016
      1 many	sexual
      1 many	aspects
      1 male	will
      1 male	then
      1 males	with
      1 males	that
      1 males	such
      1 males	receive
      1 males	punish
      1 male	society
      1 males	invest
      1 males	in
      1 males	I
      1 males	completely
      1 males	are
      1 male	represents
      1 Male	Punisher
      1 male	punisher
      1 male	parental
      1 male	only
      1 male	N
      1 male	investing
      1 male	his
      1 male	female
      1 male	favoured
      1 Male	Defector
      1 male	cooperators
      1 male	8
      1 making	them
      1 majority	of
      1 majority	invest
      1 maintenance	and
      1 made	by
      1 M	2012
      1 M	2000
      1 M	1997
      1 L	S
      1 lower	level
      1 love	and
      1 loss	to
      1 loses	its
      1 look	and
      1 long	term
      1 long	periods
      1 long	hidden
      1 logo	Issues
      1 ll	HAUERT
      1 literatures	on
      1 literatures	in
      1 listed	in
      1 line	graphs
      1 linear	form
      1 limit	of
      1 limitations	To
      1 limitations	Claussen
      1 likely	to
      1 life	with
      1 life	or
      1 life	long
      1 life	given
      1 level	where
      1 levels	Alexander
      1 level	of
      1 level	males
      1 level	in
      1 level	females
      1 level	altruism
      1 level	A
      1 less	than
      1 less	population
      1 left	x
      1 led	to
      1 law	enforcement
      1 lattice	models
      1 later	section
      1 later	period
      1 larger	than
      1 larger	difference
      1 LANGTON	C
      1 Langton	1986
      1 labeled	on
      1 L	A
      1 L	2010
      1 L	2002
      1 L	1971
      1 K	Zaballa
      1 Komdeur	J
      1 Koella	s
      1 KOELLA	J
      1 kin	altruism
      1 Ke	Zeng
      1 keywords	Introduction
      1 Keywords	Altruistic
      1 key	factor
      1 keep	the
      1 K	2007
      1 just	MD
      1 justice	still
      1 justice	religion
      1 justice	and
      1 jtbi	2000
      1 J	Pogány
      1 Jia	Y
      1 J	Grutter
      1 j	evolhumbehav
      1 J	Cosmides
      1 j	biosystems
      1 Javascript	The
      1 Javascript	is
      1 JASSS	logo
      1 JASSS	Home
      1 JASSS	Contact
      1 jasss	3058
      1 jasss	2780
      1 JASSS	2016
      1 jasss	1805
      1 JAFFE	K
      1 Jaffe	and
      1 J	1990
      1 It	would
      1 its	power
      1 its	capacity
      1 I	To
      1 It	is
      1 I	takes
      1 is	δ
      1 is	usually
      1 is	understood
      1 is	treated
      1 is	taken
      1 Issues	Issues
      1 Issues	In
      1 issues	For
      1 issue	Back
      1 is	somehow
      1 is	quite
      1 is	prob
      1 isolated	society
      1 isolated	population
      1 is	now
      1 is	notable
      1 is	no
      1 is	neglected
      1 is	necessary
      1 is	male
      1 is	likely
      1 is	introduced
      1 is	inconsistent
      1 is	in
      1 is	how
      1 is	highly
      1 I	She
      1 is	he
      1 is	hard
      1 is	given
      1 is	generally
      1 is	F
      1 is	expressed
      1 is	enhanced
      1 is	employed
      1 is	egoistic
      1 is	disabled
      1 is	defined
      1 is	considered
      1 is	conducted
      1 is	completed
      1 is	clear
      1 is	calculated
      1 is	biologically
      1 is	beyond
      1 is	also
      1 is	0
      1 I	regardless
      1 I	of
      1 in	your
      1 in	Ye
      1 in	ways
      1 involving	no
      1 involving	alternations
      1 involved	in
      1 invests	I
      1 invests	1
      1 investment	widetilde
      1 investments	in
      1 investments	are
      1 investment	remains
      1 investment	is
      1 investment	incurs
      1 investment	i
      1 Investment	Distribution
      1 investment	distribution
      1 investment	can
      1 investment	are
      1 invest	in
      1 investigated	in
      1 invest	I
      1 invest	between
      1 invest	about
      1 invasion	of
      1 in	variable
      1 invade	the
      1 in	two
      1 In	turn
      1 Introduction	Sexuality
      1 introduce	the
      1 introduced	to
      1 introduced	the
      1 into	three
      1 into	four
      1 into	account
      1 in	these
      1 inter	sexual
      1 interplay	between
      1 International	journal
      1 interest	to
      1 interest	Bird
      1 interaction	between
      1 intensity	Bulletin
      1 integrated	framework
      1 in	Structured
      1 in	structured
      1 in	steps
      1 instead	of
      1 instance	e
      1 inspire	long
      1 In	simulations
      1 in	results
      1 in	repeated
      1 in	relation
      1 in	public
      1 in	psychological
      1 in	primitive
      1 In	press
      1 in	population
      1 in	point
      1 in	penduline
      1 in	parameters
      1 In	our
      1 I	Note
      1 in	offspring
      1 in	next
      1 in	nearly
      1 in	Moran
      1 In	modern
      1 in	MATLAB
      1 in	Martu
      1 in	males
      1 initial	settings
      1 initial	proportion
      1 initial	periods
      1 initially	individual
      1 Initially	altruistic
      1 initially	A
      1 Initialize	Read
      1 in	investment
      1 in	humans
      1 in	human
      1 In	Houchmandzadeh
      1 in	his
      1 inhibition	against
      1 in	genetics
      1 infrastructures	for
      1 influence	the
      1 in	fish
      1 In	Figure
      1 in	every
      1 in	equation
      1 inequality	between
      1 in	education
      1 in	economic
      1 In	each
      1 in	each
      1 in	dynamic
      1 induced	by
      1 individual	while
      1 Individual	Variation
      1 individual	to
      1 individuals	with
      1 individuals	whose
      1 individuals	We
      1 individuals	can
      1 individuals	as
      1 individuals	are
      1 individual	provide
      1 individual	P
      1 individual	of
      1 individual	Mutant
      1 individual	equally
      1 individual	are
      1 individual	and
      1 indicators	of
      1 indicate	that
      1 indicates	the
      1 indicates	that
      1 incurs	a
      1 incurred	by
      1 incur	a
      1 in	creating
      1 increasing	the
      1 increasing	population
      1 increasing	function
      1 increase	their
      1 increases	the
      1 increases	from
      1 in	constant
      1 inconsistent	in
      1 in	computer
      1 in	competition
      1 Inclusive	fitness
      1 inclusive	fitness
      1 including	MP
      1 in	civilisations
      1 incentives	for
      1 in	C1
      1 In	C
      1 in	C
      1 in	both
      1 In	biology
      1 in	B3
      1 In	B1
      1 in	B1
      1 In	B
      1 in	B
      1 in	asexual
      1 in	artificial
      1 In	all
      1 in	all
      1 in	A
      1 in	5
      1 I	N
      1 impose	an
      1 important	problem
      1 important	parameters
      1 implicit	assumption
      1 illustrates	the
      1 illustrate	in
      1 I_i	N
      1 ignore	the
      1 ignorance	of
      1 I	gamma
      1 if	sexual
      1 I	frac
      1 I	For
      1 I	follows
      1 I	Figure
      1 if	his
      1 if	altruistic
      1 identical	normal
      1 identical	mutation
      1 identical	in
      1 I	3
      1 I	2
      1 I	1
      1 hypothesis	that
      1 hypothesis	into
      1 hunting	Evolution
      1 human	sympathy
      1 human	societies
      1 human	sociality
      1 humans	Nature
      1 human	motivation
      1 human	beings
      1 human	behavior
      1 html	KOELLA
      1 html	DOI
      1 H	Tan
      1 H	Smith
      1 How	to
      1 how	to
      1 how	sexuality
      1 However	to
      1 However	taken
      1 However	MP
      1 However	in
      1 However	as
      1 Houchmandzdeh	s
      1 HOUCHMANDZADEH	B
      1 honour	of
      1 honest	signal
      1 Home	19
      1 H	Nowak
      1 historical	observations
      1 his	own
      1 his	investment
      1 his	her
      1 him	and
      1 high	mean
      1 highly	related
      1 high	level
      1 highest	honour
      1 higher	mean
      1 higher	invasion
      1 hierarchy	of
      1 hidden	blessing
      1 her	probability
      1 heroines	with
      1 her	i
      1 her	gender
      1 hence	bears
      1 help	Preston
      1 he	benefits
      1 health	condition
      1 having	I
      1 have	to
      1 have	rarely
      1 have	not
      1 have	less
      1 have	led
      1 have	investigated
      1 have	done
      1 have	demonstrated
      1 have	broadly
      1 have	attempted
      1 have	amplifying
      1 HAUERT	C
      1 has	an
      1 Harvard	University
      1 Harmonic	Signature
      1 hard	to
      1 hard	resource
      1 happens	when
      1 HAMILTON	W
      1 Hamilton	W
      1 Hamilton	Genetics
      1 Hales	D
      1 Hales	2015
      1 H	2012
      1 H	2000
      1 Grutter	A
      1 growth	in
      1 Group	selection
      1 group	selection
      1 group	Raihani
      1 greatly	simplifies
      1 grateful	to
      1 graphs	of
      1 Grant	number
      1 g	period
      1 goods	The
      1 goods	In
      1 goods	equally
      1 goods	due
      1 goods	altruism
      1 good	genes
      1 g	lattice
      1 Given	the
      1 given	The
      1 given	the
      1 given	public
      1 given	probability
      1 given	other
      1 Gintis	Smith
      1 Gintis	et
      1 Gintis	2000
      1 G	F
      1 genetics	sexuality
      1 genetics	Mathematical
      1 genetics	argued
      1 Genetics	176
      1 Genetic	quality
      1 genetic	quality
      1 genetic	or
      1 genetical	evolution
      1 genes	Preston
      1 genes	Molecular
      1 genes	and
      1 generation	the
      1 generations	is
      1 generation	In
      1 generate	a
      1 generally	conducted
      1 generalised	to
      1 generalised	perspective
      1 generalised	Moran
      1 gender	with
      1 genders	To
      1 genders	The
      1 genders	For
      1 genders	because
      1 genders	Additionally
      1 gender	In
      1 gender	differences
      1 Gaussian	fluctuations
      1 gap	between
      1 gamma	left
      1 game	theory
      1 games	as
      1 game	Journal
      1 game	is
      1 game	In
      1 game	defined
      1 Game	A
      1 gains	the
      1 Gächter	S
      1 Gächter	2002
      1 G	2012
      1 G	1986
      1 Fund	China
      1 fundamental	difference
      1 functions	as
      1 function	in
      1 function	better
      1 fulfilled	only
      1 F	T
      1 from	Traulsen
      1 from	three
      1 from	third
      1 from	public
      1 from	nearly
      1 from	Koella
      1 from	finite
      1 from	females
      1 from	different
      1 from	Darwin
      1 from	B1
      1 from	a
      1 from	0
      1 frequency	about
      1 freedom	to
      1 framework	for
      1 frac	sum
      1 frac	N_
      1 frac	left
      1 found	to
      1 found	that
      1 found	in
      1 for	Variable
      1 For	useful
      1 for	theoretical
      1 for	their
      1 for	social
      1 For	simplicity
      1 for	simplicity
      1 for	sexuality
      1 for	sexual
      1 for	selfless
      1 for	s
      1 for	periods
      1 for	N
      1 formula	in
      1 forms	of
      1 forms	even
      1 for	moral
      1 formation	of
      1 For	male
      1 for	limited
      1 for	individual
      1 for	human
      1 for	good
      1 For	females
      1 for	females
      1 for	F
      1 For	example
      1 for	each
      1 for	different
      1 force	to
      1 forces	for
      1 force	of
      1 force	for
      1 for	attractiveness
      1 for	assistance
      1 for	A
      1 follows	the
      1 follows	MP
      1 follows	a
      1 following	Houchmandzdeh
      1 follow	an
      1 fluctuations	arising
      1 fluctuations	are
      1 Fluctuation	driven
      1 fixed	population
      1 fixation	of
      1 fitted	simulation
      1 fits	reality
      1 fitness	theory
      1 fitness	The
      1 fitness	Mate
      1 fitness	is
      1 fitness	i
      1 fitness	Hamilton
      1 fitness	for
      1 fitness	End
      1 fitness	does
      1 fitness	comes
      1 fitness	by
      1 fitness	Bell
      1 fish	Science
      1 finite	populations
      1 final	section
      1 Final	population
      1 Figure	Punishment
      1 Figure	Population
      1 figure	Figure
      1 Figure	4
      1 Figure	3indicates
      1 Figure	3
      1 Figure	2C
      1 Figure	1B
      1 fields	of
      1 fertilization	success
      1 Feminists	may
      1 female	with
      1 female	then
      1 females	show
      1 females	respectively
      1 female	society
      1 females	is
      1 females	I
      1 females	especially
      1 females	can
      1 females	both
      1 females	and
      1 female	s
      1 female	punishers
      1 female	Period
      1 Female	Defector
      1 Female	Cooperator
      1 female	cooperator
      1 Fehr	Gächter
      1 FEHR	E
      1 Feb	2016
      1 FD	will
      1 FD	This
      1 FD	right
      1 FD	respectively
      1 FD	N
      1 F	Ding
      1 FD	in
      1 FD	for
      1 FD	Female
      1 FD	Every
      1 f	denotes
      1 FD	coexist
      1 FD	beta
      1 FC	which
      1 FC	to
      1 FC	take
      1 FC	s
      1 FC	right
      1 FC	It
      1 FC	Female
      1 FC	beta
      1 FC	at
      1 FC	also
      1 FC	1
      1 Fawcett	T
      1 favour	to
      1 favours	the
      1 favours	male
      1 favoured	by
      1 favour	and
      1 fact	that
      1 factor	in
      1 factor	for
      1 F	2007
      1 extract	the
      1 extinction	of
      1 extinction	after
      1 extend	the
      1 extending	the
      1 expressed	by
      1 expressed	as
      1 express	a
      1 exponential	form
      1 exploring	the
      1 explain	many
      1 experiments	Fehr
      1 expected	payoffs
      1 expand	the
      1 exist	We
      1 exists	Yet
      1 exist	see
      1 existing	evolutionary
      1 exchange	among
      1 except	the
      1 example	of
      1 example	in
      1 examines	the
      1 exact	timing
      1 Exact	results
      1 E	Wilson
      1 evolves	as
      1 evolution	via
      1 evolution	Springer
      1 evolutions	Based
      1 evolution	is
      1 evolution	Finally
      1 evolutionary	way
      1 evolutionary	simulations
      1 evolutionary	response
      1 evolutionary	perspectives
      1 evolutionary	paths
      1 evolutionary	path
      1 evolutionary	Moran
      1 evolutionary	games
      1 evolutionary	dynamics
      1 evolutionarily	stable
      1 Evolution	and
      1 evolution	and
      1 evolhumbehav	2011
      1 evidence	of
      1 evidence	for
      1 every	traditional
      1 everyone	invests
      1 every	MP
      1 every	males
      1 Every	individual
      1 even	uninvolved
      1 even	more
      1 evenly	distributed
      1 even	if
      1 Even	at
      1 even	adoration
      1 E	Székely
      1 establishment	of
      1 established	where
      1 especially	to
      1 especially	in
      1 escape	the
      1 equivalently	sexual
      1 equilibrium	of
      1 equilibrium	from
      1 equilibrium	Discussion
      1 equation	6
      1 equal	to
      1 equals	the
      1 equally	while
      1 equally	shares
      1 e	period
      1 environment	and
      1 entire	society
      1 ensure	the
      1 enhance	the
      1 enhanced	in
      1 enforcement	and
      1 End	Set
      1 End	References
      1 employed	Surprisingly
      1 employed	B2
      1 employ	both
      1 empirical	studies
      1 elucidate	an
      1 eLS	doi
      1 egoists	Proceedings
      1 egoists	is
      1 egoists	become
      1 egoistic	contributes
      1 egoistic	because
      1 egoist	However
      1 egoism	leads
      1 E	Gächter
      1 effect	to
      1 effects	on
      1 effect	on
      1 e	F
      1 e	f
      1 education	as
      1 education	and
      1 edition	31
      1 e	defectors
      1 Ecuadorian	Amazonia
      1 economists	have
      1 Economics	Zhejiang
      1 economics	It
      1 economics	Houchmandzadeh
      1 economic	experiments
      1 Ecology	14
      1 ecological	selection
      1 e	c
      1 e	being
      1 early	period
      1 e	altruism
      1 each	of
      1 each	newborn
      1 each	generation
      1 each	figure
      1 e	a
      1 E	71
      1 E	2005
      1 Dynamics	x
      1 dynamics	Variable
      1 dynamics	raises
      1 dynamics	process
      1 dynamics	is
      1 dynamics	Figure
      1 dynamic	process
      1 dynamic	model
      1 dynamic	evolution
      1 D	we
      1 D	W
      1 during	each
      1 DUGATKIN	L
      1 Dugatkin	2007
      1 D	Thus
      1 D	the
      1 D	that
      1 D	share
      1 D	separate
      1 D	S
      1 driving	forces
      1 driven	fixation
      1 d	represents
      1 D	reach
      1 D	Pop
      1 D	Pitcher
      1 Download	PDF
      1 done	in
      1 done	based
      1 D	once
      1 dominate	the
      1 dominates	F
      1 dominated	society
      1 dominant	types
      1 dominant	over
      1 dominance	of
      1 DOI	10
      1 D	nonlinear
      1 D	N
      1 D	M
      1 division	between
      1 distribution	with
      1 distribution	was
      1 distribution	is
      1 distribution	initially
      1 Distribution	at
      1 distribution	and
      1 distributed	between
      1 Displays	a
      1 disfavours	the
      1 disfavour	parental
      1 discuss	the
      1 Discussion	of
      1 Discussion	In
      1 discussed	by
      1 disciplines	Clavien
      1 disabled	in
      1 d	is
      1 direct	rewards
      1 directly	via
      1 directly	sympathize
      1 directly	by
      1 directions	in
      1 Ding	M
      1 D	In
      1 D	in
      1 diminishing	effect
      1 dilemma	The
      1 dilemmas	International
      1 Dilemma	Other
      1 dilemma	of
      1 dilemma	in
      1 dilemma	Dawes
      1 dilemma	2007
      1 DIJK	R
      1 Dijk	et
      1 Different	Strengths
      1 different	perspectives
      1 Different	Periods
      1 Different	mutation
      1 differently	in
      1 different	levels
      1 different	evolutionary
      1 different	and
      1 difference	in
      1 deviation	of
      1 develop	a
      1 determining	the
      1 Determine	the
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      1 describing	the
      1 described	before
      1 described	as
      1 dependency	of
      1 deny	the
      1 denoted	as
      1 Deng	for
      1 d	End
      1 demonstrated	in
      1 Demon	of
      1 deleterious	mutant
      1 defined	by
      1 defined	as
      1 Defector	The
      1 defectors	to
      1 defectors	respectively
      1 defectors	regardless
      1 Defectors	initially
      1 defectors	In
      1 defectors	Figure
      1 defectors	do
      1 defectors	directly
      1 defectors	Based
      1 defectors	at
      1 Defectors	A
      1 defector	or
      1 Defector	FC
      1 defector	dominated
      1 defector	and
      1 default	in
      1 decreasing	the
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      1 decentralised	from
      1 death	Determine
      1 death	d
      1 dead	in
      1 d	c
      1 DAWES	R
      1 Dawes	Messick
      1 Darwin	to
      1 Darwinian	Demon
      1 D	2015
      1 D	2013
      1 D	1987
      1 D	1981
      1 D	1964
      1 c	αI
      1 Current	simulation
      1 Current	issue
      1 Current	directions
      1 culture	yet
      1 cultural	evidence
      1 C	The
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      1 Cross	cultural
      1 critical	evidence
      1 c	represents
      1 creating	of
      1 C	overlap
      1 could	randomly
      1 could	opt
      1 cost	to
      1 costs	the
      1 cost	of
      1 costly	signalling
      1 Costly	Signaling
      1 costly	signaling
      1 costly	commitment
      1 costly	but
      1 costly	altruistic
      1 cost	incurred
      1 cost	However
      1 cost	Houchmandzadeh
      1 cost	due
      1 cost	c
      1 cost	analysis
      1 Cosmides	L
      1 corresponding	effects
      1 Copyright	Statistics
      1 Copyright	JASSS
      1 coordinate	on
      1 cooperators	which
      1 cooperators	under
      1 cooperators	punishers
      1 cooperators	or
      1 cooperators	non
      1 cooperators	in
      1 cooperators	happens
      1 cooperator	shapes
      1 cooperators	contribution
      1 cooperators	clarified
      1 cooperators	can
      1 cooperators	as
      1 cooperator	s
      1 cooperator	is
      1 Cooperator	FD
      1 cooperator	do
      1 cooperator	defector
      1 cooperative	than
      1 cooperative	hunting
      1 cooperative	female
      1 cooperative	behavior
      1 Co	operative
      1 cooperation	researchers
      1 cooperation	proposed
      1 cooperation	Phil
      1 cooperation	might
      1 Cooperation	Journal
      1 cooperation	in
      1 cooperation	have
      1 cooperation	exist
      1 Cooperation	Basic
      1 cooperation	and
      1 contributive	and
      1 contribution	to
      1 contribution	just
      1 contribution	from
      1 contribution	by
      1 contributes	nothing
      1 contribute	Punishers
      1 contingent	upon
      1 Contact	JASSS
      1 consumption	of
      1 constraint	to
      1 constraint	in
      1 constant	scale
      1 constant	parameters
      1 constant	during
      1 constant	and
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      1 considered	altruistic
      1 Consider	an
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      1 condition	instead
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      1 concurrent	participation
      1 conclusion	withstands
      1 conclusion	is
      1 concerning	human
      1 concepts	of
      1 completely	overlap
      1 complete	for
      1 completed	through
      1 competition	with
      1 compete	for
      1 compensation	to
      1 compensation	for
      1 compensate	the
      1 compatible	genes
      1 comparison	of
      1 compare	the
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      1 community	than
      1 common	tragedy
      1 common	default
      1 commitment	to
      1 comes	from
      1 comes	at
      1 column	of
      1 colour	size
      1 collective	action
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      1 cohesion	where
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      1 cohesion	Journal
      1 cohesion	directly
      1 cohesion	because
      1 cohesion	achieved
      1 cognitive	adaptations
      1 cogent	mechanism
      1 coexist	The
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      1 co	evolution
      1 coercion	the
      1 codes	__________________________________________________________________
      1 code	for
      1 C	N
      1 clear	that
      1 clear	but
      1 Clavien	Chapuisat
      1 CLAVIEN	C
      1 Claussen	Traulsen
      1 CLAUSSEN	J
      1 classify	the
      1 classified	into
      1 clarify	that
      1 clarify	note
      1 clarified	that
      1 civilisations	Feminists
      1 civilisations	altruistic
      1 cited	Other
      1 citation	Received
      1 C	is
      1 C	in
      1 Chuankai	Cheng
      1 choose	Proceedings
      1 choice	Neff
      1 China	Other
      1 China	Grant
      1 China	bCollege
      1 Chen	Y
      1 Cheng	Ke
      1 Chenfeng	Tu
      1 Chen	Chuankai
      1 Check	We
      1 checks	on
      1 Chapuisat	M
      1 Chapuisat	2012
      1 changing	the
      1 change	of
      1 changed	the
      1 C	gains
      1 C	G
      1 certain	distribution
      1 cements	social
      1 cellular	automata
      1 C	dominates
      1 c	d
      1 C	Chapuisat
      1 causality	which
      1 case	of
      1 case	in
      1 case	for
      1 carrying	capacities
      1 care	Van
      1 care	Psychological
      1 care	praise
      1 care	in
      1 care	even
      1 care	comes
      1 capital	while
      1 capacity	The
      1 capacity	the
      1 capacity	than
      1 capacity	for
      1 capacities	of
      1 can	widen
      1 can	survive
      1 can	punish
      1 can	pay
      1 can	overcome
      1 cannot	compensate
      1 can	inspire
      1 can	influence
      1 can	escape
      1 can	choose
      1 can	benefit
      1 Cambridge	philosophical
      1 called	cooperative
      1 called	common
      1 calculations	for
      1 Calculate	payoff
      1 Calculate	fitness
      1 calculated	by
      1 C3	Simulation
      1 C3	are
      1 C3	0
      1 C2	Simulation
      1 C2	and
      1 C	2000
      1 C	200
      1 C2	0
      1 C	2
      1 C1	Simulation
      1 C1	C2
      1 C1	0
      1 C	1
      1 by	virtue
      1 by	these
      1 By	the
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      1 by	social
      1 by	pioneers
      1 by	not
      1 by	moral
      1 by	many
      1 by	Koella
      1 by	increasing
      1 by	f
      1 by	cooperators
      1 by	care
      1 by	both
      1 By	assuming
      1 by	altruistic
      1 by	a
      1 but	without
      1 but	the
      1 but	honest
      1 but	at
      1 Business	Media
      1 Bulletin	of
      1 bulletin	139
      1 B	The
      1 B	show
      1 Bshary	R
      1 B	Scelza
      1 browser	This
      1 broadly	studied
      1 b	represents
      1 breadth	of
      1 Brandt	H
      1 Boyd	Mathew
      1 Boyd	and
      1 Bowles	S
      1 both	sexual
      1 both	MP
      1 both	male
      1 both	genders
      1 both	follow
      1 both	C
      1 Book	AXELROD
      1 B	most
      1 Blessing	of
      1 blessing	of
      1 birth	rate
      1 birth	In
      1 BIRD	R
      1 Bird	et
      1 Bird	D
      1 biosystems	2014
      1 BioSystems	127
      1 biology	suggest
      1 biology	of
      1 biology	altruism
      1 biology	82
      1 biology	70
      1 biology	7
      1 biology	35
      1 biology	213
      1 biology	206
      1 biology	145
      1 biologists	have
      1 biologists	and
      1 Biological	Sciences
      1 biologically	egoistic
      1 biological	evolutionary
      1 B	In
      1 beyond	the
      1 between	Ye
      1 between	the
      1 between	reproductive
      1 between	N
      1 between	individual
      1 between	genders
      1 between	four
      1 between	cooperators
      1 between	asexual
      1 between	50
      1 between	0
      1 better	with
      1 better	Extension
      1 Besides	theoretical
      1 be	scaled
      1 be	reinforced
      1 be	qualified
      1 be	proportional
      1 be	performed
      1 be	one
      1 benefits	the
      1 benefits	others
      1 benefit	cost
      1 be	N
      1 be	more
      1 BELL	G
      1 Bell	2012
      1 believe	that
      1 beings	to
      1 behind	the
      1 behaviour	we
      1 behaviour	The
      1 behaviour	the
      1 behaviour	that
      1 behaviours	Miller
      1 behaviours	from
      1 behaviours	As
      1 behaviours	are
      1 behaviour	of
      1 behaviour	and
      1 behavior	Journal
      1 behavior	Harvard
      1 behavior	BioSystems
      1 Behavior	A
      1 behavior	33
      1 be	generalised
      1 be	fulfilled
      1 before	Population
      1 before	and
      1 before	50
      1 be	expressed
      1 be	evenly
      1 be	established
      1 be	equal
      1 been	viewed
      1 been	studied
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      1 be	detected
      1 become	dominant
      1 because	religions
      1 because	only
      1 because	at
      1 bears	no
      1 bear	a
      1 B	D
      1 bCollege	of
      1 B	C
      1 B	Bird
      1 B	Biological
      1 basic	hypothesis
      1 Basic	Book
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      1 Baseline	model
      1 based	evolutionary
      1 based	behaviours
      1 b	aSchool
      1 BARTA	Z
      1 Barta	2016
      1 balance	of
      1 Back	issues
      1 B	a
      1 b	a
      1 B3	to
      1 B3	the
      1 B3	In
      1 B3	compared
      1 B	371
      1 B2	due
      1 B	2015
      1 B	200
      1 B2	0
      1 B1	to
      1 B1	the
      1 B1	B2
      1 B1	0
      1 Axelrod	1984
      1 a	whole
      1 a	well
      1 average	value
      1 automata	Physica
      1 authors	Journal
      1 authors	How
      1 authors	For
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      1 a	unique
      1 a	typical
      1 attractive	virtue
      1 attractive	trait
      1 attractiveness	to
      1 Attractiveness	The
      1 attractiveness	the
      1 Attractiveness	Social
      1 Attractivenesss	A
      1 Attractiveness	Pop
      1 attractiveness	permit
      1 attractiveness	or
      1 attractiveness	on
      1 attractiveness	of
      1 attractiveness	For
      1 attractiveness	exist
      1 attractiveness	changed
      1 attractiveness	can
      1 attractiveness	at
      1 attractiveness	as
      1 attractiveness	are
      1 Attractiveness	and
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      1 a	tradition
      1 at	Period
      1 at	later
      1 A	The
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      1 at	capacity
      1 at	50
      1 as	x
      1 as	we
      1 A	strict
      1 a	stabilizer
      1 as	superheroes
      1 assumption	will
      1 assumption	we
      1 assumption	to
      1 assumptions	on
      1 assumption	Simulation
      1 assumptions	concerning
      1 assumptions	above
      1 Assumptions	A1
      1 assumption	is
      1 assumption	In
      1 assumption	in
      1 Assumption	Consider
      1 assuming	constant
      1 as	substructure
      1 associated	with
      1 assistance	in
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      1 As	recent
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      1 a	spatial
      1 as	P_
      1 as	other
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      1 a	society
      1 a	social
      1 as	MD
      1 A	smaller
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      1 a	single
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      1 a	significant
      1 A	Sigmund
      1 A	Shoresh
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      1 asexual	reproduction
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      1 asexual	evolutions
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      1 as	described
      1 as	defined
      1 as	compensation
      1 as	colour
      1 aSchool	of
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      1 A	S
      1 artificial	societies
      1 artificial	molecular
      1 artificial	human
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      1 are	adopted
      1 a	random
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      1 a	probability
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      1 approach	it
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      1 __________________________________________________________________	Appendix
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      1 apparent	absence
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      1 a	partial
      1 A	P
      1 any	MP
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      1 an	understanding
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      1 another	individual
      1 anonymous	referees
      1 an	integrated
      1 an	important
      1 an	identical
      1 a	new
      1 an	evolutionary
      1 and	ω
      1 and	κ
      1 and	Zaballa
      1 and	various
      1 and	this
      1 and	sympathy
      1 and	super
      1 and	state
      1 and	standard
      1 and	simulations
      1 and	signaling
      1 and	serves
      1 and	salient
      1 and	s
      1 and	routines
      1 and	Richerson
      1 and	remains
      1 and	reciprocal
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      1 and	receive
      1 and	punishment
      1 and	punisher
      1 and	psychology
      1 and	provided
      1 and	promote
      1 and	politics
      1 and	P
      1 and	other
      1 and	noted
      1 and	most
      1 and	more
      1 and	moral
      1 and	male
      1 and	law
      1 and	initial
      1 and	in
      1 and	if
      1 and	identical
      1 and	hence
      1 and	health
      1 and	Hang
      1 and	group
      1 and	function
      1 and	for
      1 and	Figure
      1 and	female
      1 and	evolution
      1 and	empirical
      1 and	egoist
      1 and	economists
      1 and	disfavours
      1 and	discussed
      1 and	d
      1 and	costs
      1 and	cooperators
      1 and	Cooperation
      1 and	compatible
      1 and	C3
      1 and	bear
      1 and	b
      1 and	authorities
      1 and	anonymous
      1 and	altruistic
      1 and	altruism
      1 and	age
      1 and	A3
      1 and	a
      1 and	5
      1 and	2
      1 and	1C
      1 and	1B
      1 and	100
      1 and	1
      1 an	asexual
      1 A	narrow
      1 an	analogous
      1 an	amplifying
      1 an	amplification
      1 Analytical	results
      1 Analysis	eLS
      1 an	altruistic
      1 analogous	to
      1 an	advantageous
      1 an	abbreviation
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      1 amplifier	for
      1 amplification	factor
      1 a	more
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      1 a	member
      1 a	measure
      1 Amazonia	Proceedings
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      1 altruist	in
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      1 altruistic	than
      1 altruistic	She
      1 Altruistic	Punishment
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      1 altruism	we
      1 altruism	versus
      1 altruism	Trivers
      1 altruism	To
      1 altruism	through
      1 altruism	The
      1 altruism	than
      1 altruism	she
      1 altruism	Quarterly
      1 altruism	Punishment
      1 altruism	on
      1 altruism	Multilevel
      1 altruism	Moreover
      1 altruism	may
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      1 altruism	involving
      1 altruism	I
      1 altruism	he
      1 altruism	Hamilton
      1 altruism	from
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      1 altruism	evolves
      1 altruism	equilibrium
      1 Altruism	Displays
      1 altruism	by
      1 altruism	but
      1 altruism	Axelrod
      1 altruism	are
      1 altruism	according
      1 Altruism	A
      1 although	MP
      1 alternative	evolutionarily
      1 alternations	of
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      1 also	shape
      1 also	reinforce
      1 also	referred
      1 also	punishes
      1 also	present
      1 also	invests
      1 also	admires
      1 a	lower
      1 ALONZO	S
      1 Alonzo	2012
      1 alone	they
      1 all	three
      1 All	the
      1 all	the
      1 a	life
      1 Algorithm	pseudo
      1 ALEXANDER	R
      1 Alexander	1987
      1 a	larger
      1 al	2012
      1 al	2010
      1 al	2008
      1 al	2007
      1 al	2002
      1 al	2001
      1 a	key
      1 A	In
      1 a	high
      1 a	Harmonic
      1 a	given
      1 agents	Shutters
      1 agent	in
      1 a	gender
      1 age	can
      1 against	norm
      1 again	show
      1 after	some
      1 African	journal
      1 A	Figure
      1 Á	Fawcett
      1 advertise	the
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      1 advantageous	in
      1 a	driving
      1 adoration	and
      1 adopted	from
      1 adopt	a
      1 admires	MP
      1 admire	female
      1 admire	any
      1 admire	according
      1 admiration	Ye
      1 admiration	Miller
      1 admiration	if
      1 adjusted	models
      1 adjusted	model
      1 adjusted	for
      1 a	diminishing
      1 a	different
      1 a	deleterious
      1 a	defector
      1 Additionally	we
      1 Additionally	robust
      1 Additionally	altruistic
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      1 Actually	females
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      1 actor	s
      1 action	The
      1 A	costly
      1 a	cooperator
      1 A	cooperative
      1 a	comparison
      1 a	common
      1 a	cogent
      1 Acknowledgements	This
      1 __________________________________________________________________	Acknowledgements
      1 acknowledge	Chenfeng
      1 a	certain
      1 accuracy	of
      1 account	Even
      1 Accepted	24
      1 a	case
      1 academy	of
      1 abundant	public
      1 Abstract	Current
      1 absence	of
      1 A	Brandt
      1 A	Bowles
      1 above	The
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      1 About	JASSS
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      1 about	0
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      1 A3	C3
      1 A2	C2
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      1 A	2008
      1 A	2007
      1 A	2005
      1 A	200
      1 A2	0
      1 A1	C1
      1 A1	A2
      1 A1	0
      1 a0003442	pub2
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      1 9	Figure
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      1 9904	992
      1 9865	200
      1 9780470015902	a0003442
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      1 8	where
      1 8	suffers
      1 8	Mean
      1 8	Figure
      1 8	examines
      1 8	C
      1 8	because
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      1 8	A
      1 8721	00173
      1 830	C3
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      1 80113	4
      1 7	where
     17 the	society
      1 7	Simulation
      1 7	ll
      1 7	In
      1 7	and
      1 79	82
      1 78	doi
      1 71	doi
      1 71	2
      1 70	5
      1 6	x
     16 value	of
      1 6	The
      1 6	Simulation
      1 6	shows
      1 6868	137
      1 66	doi
      1 6	5
      1 64	78
      1 6	1305
      1 6	1
      1 60	71
      1 60	66
     16 0	4
      1 6	0
      1 5	where
     15 variable	size
     15 the	evolution
      1 5	The
      1 5	the
      1 5	tests
     15 size	population
      1 5	Simulation
      1 5	shows
      1 5	respectively
     15 punishment	and
     15 and	sexual
      1 5	and
     15 altruistic	punishment
      1 5962	171
      1 5833	1905
      1 5833	1858
      1 57	doi
      1 551	B2
      1 54	1
      1 5193	05
      1 5	1410
      1 50	respectively
      1 50	and
      1 4	where
      1 4	Simulation
      1 4	Nearly
     14 in	Figure
      1 4	http
      1 4	html
     14 doi	10
      1 4	CLAUSSEN
      1 4489	1390
      1 442	C2
      1 4416	200
      1 4396	991
      1 4392	551
      1 415	6868
      1 406	PRESTON
      1 406755	VAN
     14 0	5
      1 3	Simulation
     13 sexual	selection
     13 s	0
      1 3	s
      1 3	P_
      1 3indicates	the
     13 et	al
     13 as	a
      1 394	406
      1 38	POTGIETER
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      1 35	2
      1 3501	830
      1 342	doi
      1 3	394
      1 3	331
      1 331	342
      1 33	1
      1 327	5962
      1 3208	442
      1 31	Mar
      1 3	1375
      1 31	3
      1 3058	Save
     12 that	the
      1 2	Simulation
     12 results	of
      1 2	P_
      1 2	MP
      1 2	Model
      1 2	Javascript
      1 2	individual
      1 2	illustrates
      1 2	if
     12 i	e
      1 2	Given
      1 2	For
      1 2	Figure
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      1 279	1734
      1 2	79
      1 2780	SOBER
      1 267	1456
      1 2601	200
      1 24	Feb
      1 22	1
      1 2	169
      1 213	1
      1 211	4489
      1 2111	GINTIS
      1 2	111
      1 20	When
      1 20	http
      1 20	html
      1 206	2
     12 0	6
      1 2	025101
      1 2016	Therefore
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      1 2016	Individual
      1 2016	Abstract
      1 2016	
      1 2015	where
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      1 2015	N
      1 2015	In
      1 2015	Fluctuation
      1 2015	assumption
      1 2015	and
      1 2015	Altruism
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      1 20150087	doi
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      1 2012	Selection
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      1 2010	proposed
      1 2010	Our
      1 2010	Co
      1 200	Treatments
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      1 200	B1
      1 200	A3
      1 200	A2
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      1 2008	Analytical
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      1 2007	via
      1 2007	To
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      1 2007	Sexual
      1 2007	Selection
      1 2007	Researches
      1 2007	Inclusive
      1 2007	Behavior
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      1 2005	Non
      1 2005	Miller
      1 2005	Genetic
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      1 2002	The
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      1 2002	New
      1 2002	Gintis
      1 2002	Cross
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      1 2001	Religions
      1 2001	McAndrew
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      1 2000	The
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      1 2000	2111
      1 2000	1239
      1 2	000
      1 1	x
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      1 1	P_
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      1 1	making
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      1 1	For
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     11 C	and
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      1 1997	The
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      1 1990	Sober
      1 1990	Group
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      1 1983	The
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      1 1979	1985
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      1 1964	The
      1 1964	Dugatkin
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      1 1958	Random
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      1 19	38
      1 1907	HOUCHMANDZADEH
      1 1905	1907
      1 1859	BOYD
      1 1858	1859
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      1 17	Oct
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      1 1784	1790
      1 176	3
      1 1734	1784
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      1 17	11537
      1 169	179
      1 1687	20150087
      1 1	64
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      1 1467	8721
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      1 1390	1396
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      1 1375	1380
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      1 127	60
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      1 1	200
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      1 0	9904
      1 0	9865
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     10 5	000
      1 0	4416
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      1 0	3501
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      1 000	It
      1 000	Final